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Erad control of protein quality and quantity via the ubiquitin-proteasome system (UPS) Ubiquitination is a sequential process orchestrated by core E1 ubiquitin-activating enzymes, general E2 ubiquitin-conjugating enzymes, and selective E3 ubiquitin ligases, which, in concert, conjugate ubiquitin moieties in select topologies to a substrate (Fig